Concepts regarding information infrastructure, or cyberinfrastructure, have evolved over the past thirty years as organizations and institutions developed best practices for “cyber-based information systems.”  Lincoln Stein reported in 2008 on the state of affairs of the “biological cyberinfrastructure”, describing four areas of development needed to make data and tools available to enable computational analyses and to facilitate the exchange of knowledge from those analyses.  SERNEC, the SouthEast Regional Network of Expertise and Collections, was funded through two National Science Foundation grants to mobilize the community of curators and herbarium affiliates in the Southeast USA to make herbarium specimens and associated data available to the public.  The community has been engaged in developing, maintaining and sustaining the four cyberinfrastructure areas outlined by Stein: data infrastructure, computational infrastructure, communication infrastructure and human infrastructure.  The SERNEC effort has generated 5.5 million electronic specimen records to date.  Most of the herbaria engaged in the project have fully incorporated the digitization process into their day-to-day workflows.  We have demonstrated that a tiered network of herbaria of all sizes, and the people associated with them, has tremendous potential to generate a very large high quality dataset and to drive collaborations among associated researchers.  In this presentation we report on our progress, with particular emphasis on the human infrastructure that generated the 15 million specimens housed in the 233 herbaria in the Southeast USA.  The community has embraced phases of the digitization process, including by-hand entry of skeletal data, imaging, citizen science driven label transcription and georeferencing.  The challenge of sustaining this effort across academic generations will be explored.  In addition, we are developing strategies to encourage the community to engage in extended specimen activities.  We discuss the addition of Bionomia as a tool to both connect scientists with these specimens and to extend specimen value.  We will explore strategies to engage our community to embrace Bionomia as a part of our growing cyberinfrastructure.  The addition of Bionomia to our toolkit demonstrates the organizational value of maintaining and sustaining all four aspects of a cyberinfrastructure to maximize community productivity.